Harrison, G., H.P. Dunleavy, L.F. Vasquez-Valverde, A.I. Del Pozo-Valdivia, K. Ivanov, P.E. Marek. 2024. Arthropod diversity in shallow subterranean habitats of the Appalachian Mountains. Subterranean Biology, submitted.

Mikel-Stites, M., P.E. Marek, M. Hellier, A. Staples. 2024. Left-right tympanal size asymmetry in the parasitoid fly Ormia ochracea, microPublication Biology, in review.

Menegatti, C., P. Banks, K. Knott, A.J. Briganti, S.V.G. McNally, P.E. Marek, A.M. Brown, T.H. Jones, E.Y. Mevers. 2024. Neuromodulating alkaloids from millipede defensive secretions. [ChemRxiv Preprint]

Banks, P., E.M. Funkhouser, A.M. Macias, B. Lovett, S. Meador, A. Hatch, H.M. Garraffo, K.C. Cartwright, M.T. Kasson, P.E. Marek, T.H. Jones, E.Y. Mevers. 2024. The chemistry of the defense secretions of three species of millipedes in the genus Brachycybe, Journal of Chemical Ecology, 1–11.    [Open access] [ChemRxiv Preprint]

Shear, W.A., P.E. Marek. 2024. The millipede family Striariidae Bollman, 1893. IX. The identity of Striaria californica Cook, 1899, and the new genus Bayaria for Striaria nana Loomis, 1936, with a key to genera and an annotated checklist of the Striariidae (Diplopoda, Chordeumatida, Striarioidea), Zootaxa, 5463: 524–544.    [Link]

Bergh, J.C., P.E. Marek, B.D. Short, J.H. Skevington, F. Thompson, 2023. The identity of Neocnemodon calcarata (Loew) (Diptera: Syrphidae), a specialized flower fly predator of woolly apple aphid, In: K. Jordaens, X. Mengual, J. Skevington (eds.) Contributions to Dipterology in honour of F. Christian Thompson, Studia dipterologica, Supplement 23: 147–163.    [Link]

P.E. Marek, C.L. Hall, C. Lee, J. Bailey, M. Berger, M. Kasson, W.A. Shear. 2023. A new species of Illacme from southern California (Siphonophorida, Siphonorhinidae), ZooKeys, 1167: 265–291.    [Open access]

Shear, W.A., P.E. Marek. 2023. The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea), Zootaxa, 5264: 323–340.    [Link]

Mikel-Stites, M., M.K. Salcedo, J.J. Socha, P.E. Marek, A.E. Staples. 2023. Reconsidering tympanal-acoustic interactions leads to an improved model of auditory acuity in a parasitoid fly, Bioinspiration & Biomimetics, 18 (2023) 035007.    [Open access]

P.E. Marek, Shear, W.A. 2022. Quick guide: Myriapoda, Current Biology, 32: 1294–1296.    [Link]

Shear, W.A., P.E. Marek, J.E. Bond, T. Wesener. 2022. The millipede family Striariidae Bollman, 1893. VII. Petra sierwaldae, n. gen., n. sp., a minute millipede from Idaho, USA (Diplopoda, Chordeumatida, Striariidae), Zootaxa, 5213: 287–293.    [Link]

Shear, W.A., P.E. Marek. 2022. The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea), Zootaxa, 5205: 501–531.    [Open access]

Vasquez-Valverde, L.F., P.E. Marek. 2022. Phylogenetic review of the millipede genus Cherokia Chamberlin, 1949 (Polydesmida: Xystodesmidae), ZooKeys, 1106: 141–163.    [Open access]

Hennen, D.A., J.C. Means, P.E. Marek. 2022. A revision of the wilsoni species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae), ZooKeys, 1096: 17–118.    [Open access]

Jones, T., D.P. Harrison, C. Menegatti, E. Mevers, K. Knott, P. Marek, D. Hennen, M. Kasson, A. Macias, B. Lovett, R. Saporito. 2022. Deoxybuzonamine isomers from the millipede Brachycybe lecontii (Platydesmida: Andrognathidae), Journal of Natural Products, 85: 1134–1140.   [Link]    Featured on the issue’s cover.

Shear, W.A., P.E. Marek. 2022. The milliped family Striariidae Bollman, 1893. V. Stegostriaria dulcidormus, n. gen., n. sp., Kentrostriaria ohara, n. gen., n. sp., and the convergent evolution of exaggerated metazonital crests (Diplopoda, Chordeumatida, Striarioidea). Zootaxa, 5094: 461–472.    [Open access]

Shear, W.A., P. Nosler, P.E. Marek. 2022. The millipede family Striariidae Bollman, 1893. IV. Amplaria oedipus, n. sp., with a secondary sexual modification of males unique among millipedes (Diplopoda, Chordeumatida, Striarioidea). Zootaxa, 5099: 137–145.  [Open access]

Marek, P.E., B.A. Buzatto, W.A. Shear, J.C. Means, D.G. Black, M.S. Harvey, J. Rodriguez. 2021. The first true millipede—1306 legs long. Scientific Reports, 11: 1–8.   [Open access]

Shear, W.A., P.E. Marek. 2021. Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae). Zootaxa, 4975: 81–126.    [Open access]

Means, J.C., D.A. Hennen, P.E. Marek. 2021. A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae). ZooKeys, 1030: 1-180.    [Open access]

Means, J.C., D.A. Hennen, T. Tanabe, P.E. Marek. 2021. Phylogenetic systematics of the millipede family Xystodesmidae. Insect Systematics and Diversity, 5: 1-26.    [Open access]    Featured on the issue’s cover.

Wong, V.L., D.A. Hennen, A.M. Macias, M.S. Brewer, M.T. Kasson, P.E. Marek. 2020. Natural history of the social millipede Brachycybe lecontii Wood, 1864. Biodiversity Data Journal, 8: e50770.    [Open access]

Wong, V.L., P.E. Marek. 2020. Structure and pigment make the eyed elater’s eyespots black. PeerJ, 8: 8161.    [Open access] [PeerJ Preprint]

Shear, W.A., P.E. Marek 2019. Urochordeumatidae Silvestri, 1909, a millipede family endemic to Washington State, USA (Chordeumatida, Striariidea, Striarioidea). Zootaxa, 4657: 352-360.    [Link]

Macias, A.M, P.E. Marek, E. M. Morrissey, M.S. Brewer, D.P.G. Short, C.M. Stauder, K.L. Wickert, M.C. Berger, A.M. Methany, J.E. Stajich, G. Boyce, R.V.M. Rio, D.G. Panaccione, V.L. Wong, T.H. Jones, M. Kasson. 2019. Diversity and function of fungi associated with the fungivorous millipede, Brachycybe lecontii. Fungal Ecology, 41: 187-197.    [Link]    [bioRxiv Preprint]

Sierwald, P., D.A. Hennen, X.J. Zahnle, S. Ware, P.E. Marek. 2019. Taxonomic synthesis of the eastern North American millipede genus Pseudopolydesmus (Diplopoda: Polydesmida: Polydesmidae), utilizing high-detail ultraviolet fluorescence imaging. Zoological Journal of the Linnean Society, 1-26.    [Open access]

Nguyen, A.D., P. Sierwald, P.E. Marek. 2019. The pill millipedes of Vietnam: a key to genera and descriptions of five new species (Diplopoda: Glomerida: Glomeridae). Raffles Bulletin of Zoology, 67: 260-297.    [Open access]

Shorter, P.L., D.A. Hennen, P.E. Marek. 2018. Cryptic diversity in Andrognathus corticarius Cope, 1869 and description of a new Andrognathus species from New Mexico (Diplopoda, Platydesmida, Andrognathidae). ZooKeys, 786: 19-41.    [Open access]

Lancaster, J., A. Khrimian, S. Young, B. Lehner, K. Luck, A. Wallingford, S. Ghosh, P. Zerbe, A. Muchlinski, P. Marek, M. Sparks, J. Tokuhisa, C. Tittiger, T. Köllnerd, D. Weber, D. Gundersen-Rindal, T. Kuhar, D. Tholl. 2018. De novo formation of an aggregation pheromone precursor by an isoprenyl diphosphate synthase-related terpene synthase in the harlequin bug. Proceedings of the National Academy of Sciences, USA.    [Open access]

Rodriguez, J., T.H. Jones, P. Sierwald, P.E. Marek, W.A. Shear, M.S. Brewer, J.E. Kocot, J.E.Bond 2018. Step-wise evolution of complex chemical defenses in millipedes: a phylogenomics approach. Scientific Reports, 8: 3209.    [Open access]

Marek, P.E., J.C. Means, D.A. Hennen. 2018. Apheloria polychroma, a new species of millipede from the Cumberland Mountains (Polydesmida: Xystodesmidae). Zootaxa, 4375: 409-425.    [Open access]

Yang, F., K. Chan, P.E. Marek, P.M. Armstrong, P. Liu, J.E. Bova, J.N. Bernick, B.E. McMillan, B.G. Weidlich, S.L. Paulson 2018. Cache Valley Virus in Aedes japonicus japonicus mosquitoes, Appalachian Region, United States. Emerging Infectious Diseases, 24: 553-557.     [Open access]

Means, J.C.** and P.E. Marek. 2017. Is geography an accurate predictor of evolutionary history in the millipede family Xystodesmidae? PeerJ, 5: e3854.    [Open access]

Shear, W.A., P. Nosler**, and P.E. Marek 2017. The identity of Amplaria nazinta (Chamberlin, 1910): a century-old millipede mystery resolved (Diplopoda, Chordeumatida, Striariidae). Zootaxa, 4311: 233-240.    [Link]

Marek, P.E. 2017. Ultraviolet-induced fluorescent imaging for millipede taxonomy. Research Ideas and Outcomes, 3: 1-14.    [Open access]

Marek, P.E., J. Krejca, and W.A. Shear. 2016. A new species of Illacme Cook & Loomis, 1928 from Sequoia National Park, California, with a world catalog of the Siphonorhinidae (Diplopoda, Siphonophorida). ZooKeys, 626: 1-43.    [Open access]

Shear, W.A., R.L. Ferreira, L.F.M. Iniesta, P.E. Marek 2016. A millipede missing link: Dobrodesmidae, a remarkable new polydesmidan millipede family from Brazil with supernumerary rings (Diplopoda, Polydesmida), and the establishment of a new suborder Dobrodesmidea. Zootaxa, 4178: 371-390.    [Link]

Dellinger, T., V. Wong**, and P.E. Marek. 2016. Makelabels: a Bash script for generating data matrix codes for collection management. Biodiversity Data Journal, 4: 1-9.    [Open access]

Garcia, A.J.**, S. Priya, and P.E. Marek. 2015. Understanding the locomotion and dynamic controls for millipedes: Part 1 – Kinematic analysis of millipede movements. Proceedings of the ASME 2015 Conference on Smart Materials, Adaptive Structures and Intelligent Systems, 1 – 10.    [Link]

Meyer, W.M., J. Eble, K. Franklin, R.B. McManus**, S.L. Brantley, J. Henkle**, P.E. Marek, W.E. Hall, C.A. Olson, R. McInroy**, E.M. Bernal Loaiza**, R.C. Brusca, and W. Moore. 2015. Ground-Dwelling Arthropod Communities of a Sky Island Mountain Range in Southeastern Arizona, USA: Obtaining a Baseline for Assessing the Effects of Climate Change. PLoS ONE, 10(9): 1-18.    [Open access]

Means, J.C.**, E.A. Francis**, A.A. Lane**, and P.E. Marek. 2015. A general methodology for collecting and preserving xystodesmid and other large millipedes for biodiversity research. Biodiversity Data Journal, 3: 1 – 17.    [Open access]

Marek, P. E., and W. Moore. 2015. Discovery of a glowing millipede in California and the gradual evolution of bioluminescence in Diplopoda. Proceedings of the National Academy of Sciences, USA. 112, 6419-6424.    [Open access]

Harris, M.C.**, E.J. Dotseth, B.T. Jackson, S.D. Zink, P.E. Marek, L.D. Kramer, S.L. Paulson, and D.M. Hawley. 2015. La Crosse virus in Aedes japonicus japonicus mosquitos in the Appalachian Region. Emerging Infectious Diseases, 21.    [Open access]

Marek, P.E., T. Tanabe, and S. Sierwald. 2014. A species catalog of the millipede family Xystodesmidae (Diplopoda: Polydesmida). Virginia Museum of Natural History Special Publications, 17: 1 – 117.    [Open access]

Kamali, M.**, A. Peery**, P.E. Marek, C. Antonio-Nkondjio, C. Ndo, Z. Tu, F. Simard, and I.V. Sharakhov. 2014. Temporal diversification of major African malaria vectors inferred from multigene phylogenetics. PLoS ONE 9(4): 1-9.    [Open access]

Marek, P.E., W.A. Shear, and J.E. Bond. 2012. A redescription of the leggiest animal, the millipede Illacme plenipes with notes on its natural history and biogeography (Diplopoda, Siphonophorida, Siphonorhinidae). ZooKeys, 241, 77–112.    [Open access]

Marek, P.E., D.R. Papaj, J. Yeager, S. Molina**, and W. Moore. 2011. Bioluminescent aposematism in millipedes. Current Biology, 21, R680-R681.    [Open access]
Video abstract online at Cell Press YouTube channel.

Marek, P.E. 2010. A revision of the Appalachian millipede genus Brachoria Chamberlin, 1939 (Polydesmida, Xystodesmidae, Apheloriini). Zoological Journal of the Linnean Society, 159, 817-889.    [Link]

Shear, W.A. and P.E. Marek. 2009. Andrognathus hoffmani, n. sp., a second species in the genus and the first species of Andrognathidae from México (Diplopoda, Platydesmida, Andrognathidae). Festschrift in honor of Richard Hoffman, Memoirs of the Virginia Museum of Natural History, 16, 149-158.    [Open access]

Marek, P.E. and J.E. Bond. 2009. A Müllerian mimicry ring in Appalachian millipedes. Proceedings of the National Academy of Sciences, USA. 106, 9755-9760.    [Open access]
Commentary by Merrill, R.M. and C.D. Jiggins. 2009. Müllerian mimicry: sharing the load reduces the legwork, Current Biology, 19, 687-689.    [Open access]

Walker, M.J.**, A.K. Stockman, P.E. Marek, and J.E. Bond. 2009. Pleistocene glacial refugia across the Appalachian Mountains and coastal plain in the millipede genus Narceus: Evidence from population genetic, phylogeographic, and paleoclimatic data. BMC Evolutionary Biology. 9, 25.    [Open access]

Marek, P.E. and J.E. Bond. 2007. A reassessment of apheloriine millipede phylogeny: additional taxa, Bayesian inference, and direct optimization. Zootaxa. 1610, 27-39.    [Link]

Marek, P.E. and J.E. Bond. 2006. Phylogenetic systematics of the colorful, cyanide-producing millipedes of Appalachia (Polydesmida, Xystodesmidae, Apheloriini) using a total evidence Bayesian approach. Molecular Phylogenetics and Evolution. 41, 704-729.    [Link]

Marek, P.E. and J.E. Bond. 2006. Biodiversity hotspots: Rediscovery of the world’s leggiest animal. Nature. 441, 707.    [Link]

Marek, P.E. and D.H. Kavanaugh. 2005. The evolutionary relationships of North American Diplous Motschulsky (Coleoptera: Carabidae: Patrobini) inferred from morphological and molecular evidence. Invertebrate Systematics. 19, 145-168.    [Link]

Marek, P.E., J.E. Bond, and P. Sierwald. 2003. Rhinocricidae Systematics II: A species catalog of the Rhinocricidae (Diplopoda: Spirobolida) with synonymies. Zootaxa. 308, 1-108.    [Open access]

Bond, J.E. and P.E. Marek. 2003. Rhinocricidae Systematics I: The taxonomic placement of the species of Zipyge Chamberlin, 1925 and Oxypygides Chamberlin, 1922 (Diplopoda: Spirobolida: Rhinocricidae: Oxypyginae). Zootaxa. 292, 1-8.    [Open access]

** student co-authored publications

Non peer-reviewed publications

Marek, P.E. 2013. Xystodesmidae, in: Wikispecies, free species directory, online at

Marek, P.E. 2012. A clever experiment, in: Creatures of Light, American Museum of Natural History iPad app, online at

Marek, P.E. 2012. Environment 4, in: Isopod: The Roly Poly Science Game, Xylem and Phloem LLC iPad app, online at

Marek, P.E. 2011. Motyxia, the Sierra luminous millipedes, in: Wikipedia, the free encyclopedia, online at

Marek, P.E. 2010. Brachoria Chamberlin 1939. The Appalachian Mimic Millipedes. Version 16 September 2010. in The Tree of Life Web Project,